This is a zoosporangium of Halophytophthora epistomium,
an oomycote that can be found regularly, though rarely, in
saltmarsh ecosystems, on leaves of plants at the marsh edge (e.g.,
live oak) that fall into marsh pools. When zoospores have matured,
the sporangium pops the cap of the dehiscence tube off, a plug is
extruded, and the zoospores swim out of the opened tube. H.
epistomium can enwrap hyphae of true fungi with its own, and
feed on the true-fungal hyphae. See Newell & Fell, 1996, Cues for
zoospore release by marine oomycotes in naturally decaying
submerged leaves, Mycologia 88:934-938.