Saltmarsh coffeebean snails (Melampus bidentatus) are
grazers of standing-decaying cordgrass, just as are
saltmarsh periwinkles , but they
apparently don't take fungal material so intensively that they
cause reductions in fungal productivity like the periwinkles can.
Coffeebean grazing actually stimulates fungal growth
(Graça, Newell & Kneib, 2000, Mar. Biol. 136:281-289).
Coffeebean snails are most obvious at the upland edges of smooth-cordgrass
marshes, but I have found small ones deep in the marsh, and they generally
hide during the day, so they may be more widely distributed than is now
generally thought. Coffeebean snails eat all sorts of marsh-edge
plant litter (marshgrass, oak leaves, palm fronds, etc.) but they
like standing-decaying cordgrass just fine. I have raised them
from 2-mm babies to full adult size (13-mm
shell length) on standing-decaying cordgrass as the only food. See Newell
& Porter, 2000, pp. 159-185 in Weinstein & Kreeger, Concepts and
Controversies in Tidal Marsh Ecology, Kluwer.