Snails aren't the only grazers of naturally-decaying
marshgrass. Amphipods do it too. These are adults of
Uhlorchestia spartinophila, the most common shoot-associated
amphipod in the smooth-cordgrass marsh. Here is one that is not anesthetized . These talitrids are almost
always found on cordgrass shoots, not down on or in the marsh
sediment. Their grazing is not as intense as that of periwinkles,
and usually only the leaf and fungal material at and near the leaf surface
is removed (as in this image). I have watched them as they carefully pick
out the parts of the decaying leaf that they want (they're not bashful
when they're hungry). They will scoop out the contents of fungal
ascomata (the sexual-spore-containing structures) in decaying
leaves, and they will eat pure mycelium of saltmarsh ascomycetes if
it is offered to them. See Graça et al, 2000, Mar Biol
136:281-289; Newell & Porter, 2000, pp. 159-185 in Weinstein
& Kreeger, Concepts and Controversies in Tidal Marsh Ecology,
Kluwer.